A New Species of Diplocheila Brulle from North America, with Notes on Female Reproductive Tract Characters in Selected Licinini and Implications for Evolution of the Subgenus Isorembus Jeannel (coleoptera: Carabidae: Licinini)

-Diplocheila (lsorembus) cross;' new species, is described, from central and southeastern United States. Genitalic and internal female reproductive tract structures for Licinini are presented for exemplars of Dicaelina, Licinina and, Lestignathina. An evagination of the bursa common to Diplocheila and Dicaelus is a synapomorphy for Dicaelina. The spennatheca and appended gland in the subgenus Isorembus provide characters that are congruent with Ball's 1959 phylogeny for the group and suggest resolution for several ambiguous relationships. Apparent transfonnations of the female reproductive tract morphology include a shift from an elongate spennathecal gland and a spennatheca with a strongly differenciated tip to a spherical spennathecal gland and more unifonn spennatheca. Other aspects of the tract appear to be highly conserved among the licinine taxa examined.